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Acknowledgements This work was supported by the National Key R&D Program of China (2017YFA0205501), the National Natural Science Foundation of China (81722024, 81571728 and 31530026), the Key Research Program of Frontier Sciences (QYZDY-SSW-SMC013), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDA09030306 and XDPB0304), the Youth Innovation Promotion Association (2014078) and the Sanming Project of Medicine in Shenzhen (SZSM201612031)